Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them. Usually the aim is correlation, demonstrating that a particular horizon in one geological section represents the same period of time as another horizon at some other section. The fossils are useful because sediments of the same age can look completely different because of local variations in the sedimentary environment. For example, one section might have been made up of clays and marls while another has more chalky limestones, but if the fossil species recorded are similar, the two sediments are likely to have been laid down at the same time.
Biostratigraphy originated in the early 19th century, where geologists recognised that the correlation of fossil assemblages between rocks of similar type but different age decreased as the difference in age increased. The method was well-established before Charles Darwin explained the mechanism behind it--evolution.
Ammonites, graptolites, archeocyathids, and trilobites are index fossils that are widely used in biostratigraphy. Microfossils such as acritarchs, chitinozoans, conodonts, dinoflagellate cysts, ostracods, pollen, spores and foraminiferans are also frequently used. Different fossils work well for sediments of different ages; trilobites, for example, are particularly useful for sediments of Cambrian age. To work well, the fossils used must be widespread geographically, so that they can occur in many different places. They must also be short lived as a species, so that the period of time during which they could be incorporated in the sediment is relatively narrow. The longer lived the species, the poorer the stratigraphic precision, so fossils that evolve rapidly, such as ammonites, are favoured over forms that evolve much more slowly, like nautiloids. Often biostratigraphic correlations are based on a fauna, not an individual species, as this allows greater precision. Further, if only one species is present in a sample, it can mean that (1) the strata were formed in the known fossil range of that organism; (2) that the fossil range of the organism was incompletely known, and the strata extend the known fossil range. For instance, the presence of the trace fossil Treptichnus pedum was used to define the base of the Cambrian period, but it has since been found in older strata.
Fossil assemblages were traditionally used to designate the duration of periods. Since a large change in fauna was required to make early stratigraphers create a new period, most of the periods we recognise today are terminated by a major extinction event or faunal turnover.
A stage is a major subdivision of strata, each systematically following the other each bearing a unique assemblage of fossils. Therefore, stages can be defined as a group of strata containing the same major fossil assemblages. French palaeontologist Alcide d'Orbigny is credited for the invention of this concept. He named stages after geographic localities with particularly good sections of rock strata that bear the characteristic fossils on which the stages are based.
In 1856 German palaeontologist Albert Oppel introduced the concept of zone (also known as biozones or Oppel zone). A zone includes strata characterised by the overlapping range of fossils. They represent the time between the appearance of species chosen at the base of the zone and the appearance of other species chosen at the base of the next succeeding zone. Oppel's zones are named after a particular distinctive fossil species, called an index fossil. Index fossils are one of the species from the assemblage of species that characterise the zone.
The zone is the fundamental biostratigraphic unit. Its thickness range from a few to hundreds of metres, and its extant range from local to worldwide. Biostratigraphic units are divided into six principal kinds of biozones:
To be useful in stratigraphic correlation index fossils should be: